You can experiment also with layer' and brush blending modes, such as Soft Light, Screen, Multiply, etc.After it's may be needed to make some areas deeper (darker) - for example, shaded areas deeply among strands near their roots - or set some light and highlight accents in the lights.So, try at first to place basic shading with a simple brush you prefer to paint usually, hard or soft, and then put the fur strands above, according to the form and lighting.
All the brushes have a minor color and lightness jitter to make the fur more realistic. Usually this underpainting should be darker than fur, as it would be in the shadow. As you know, there's a number of basic shading technics in painting and drawing, and I like this one as the most appropriate for me - I find it one of the most rational by painting the fur - to start coloring with a neutral, halftone local color, then shade it according to the volume and lighting.
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